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When you have it equipped, you slowly regain health & EVE whilst standing in or under a large leak of falling water.

The Fountain of Youth Gene Tonic, is awarded to the player in BioShock 2 upon achieving level 4 of Spider Splicer research. In the Minerva’s Den, it is found in a flooded crawlspace in the Access Shaft area, of the level The Thinker.